diff --git a/jstests/multiVersion/genericBinVersion/query-integration-timeseries/timeseries_dollar_prefixed_timefield_errors.js b/jstests/multiVersion/genericBinVersion/query-integration-timeseries/timeseries_dollar_prefixed_timefield_errors.js
index fb37c823a8c..c97d96d926b 100644
--- a/jstests/multiVersion/genericBinVersion/query-integration-timeseries/timeseries_dollar_prefixed_timefield_errors.js
+++ b/jstests/multiVersion/genericBinVersion/query-integration-timeseries/timeseries_dollar_prefixed_timefield_errors.js
@@ -136,14 +136,16 @@ function assertCollectionsQueryable(db) {
 }
 
 testPerformReplSetRollingRestart({
-    // support for $-prefixed timeField/metaField was removed in 8.3
+    // Support for $-prefixed timeField/metaField was removed in v8.3.
     startingVersion: {binVersion: "last-lts"},
     setupFn: setUp,
     beforeRestart: () => {},
-    afterSecondariesHaveRestarted: (primaryConnection, originalPrimary) => {
-        // Create on the node that wasn't restarted (still on last-lts).
-        assertCreateDollarCollectionsWorks(getDB(originalPrimary));
-        assertCollectionsQueryable(getDB(primaryConnection));
+    afterSecondariesHaveRestarted: (primaryConnection) => {
+        // Check that collections can be created and queried while secondaries are on the new
+        // version and primaries are on the old version.
+        const db = getDB(primaryConnection);
+        assertCreateDollarCollectionsWorks(db);
+        assertCollectionsQueryable(db);
     },
     afterPrimariesHaveRestarted: (primaryConnection) => {
         const db = getDB(primaryConnection);
diff --git a/jstests/multiVersion/libs/mixed_version_fixture_test.js b/jstests/multiVersion/libs/mixed_version_fixture_test.js
index d925a15f737..7283a2a6f88 100644
--- a/jstests/multiVersion/libs/mixed_version_fixture_test.js
+++ b/jstests/multiVersion/libs/mixed_version_fixture_test.js
@@ -52,6 +52,9 @@ export function testPerformReplSetRollingRestart({
             {
                 ...startingVersion,
             },
+            {
+                ...startingVersion,
+            },
         ],
         // Create a copy of the options since the callee may modify them.
         nodeOptions: copyJSON(startingNodeOptions),
@@ -76,15 +79,11 @@ export function testPerformReplSetRollingRestart({
             " and options " +
             tojsononeline(restartNodeOptions),
     );
-    // Save a reference to the node that was primary before restarting secondaries. Pass it to the
-    // callback so tests can run operations that only the old version supports; the current primary
-    // after upgradeSecondaries() may be the restarted node in a 2-node set.
-    const originalPrimary = primaryConnection;
     rst.upgradeSecondaries({...restartVersion, ...copyJSON(restartNodeOptions)});
     primaryConnection = rst.getPrimary();
 
     jsTest.log.info("Calling the afterSecondariesHaveRestarted function");
-    afterSecondariesHaveRestarted(primaryConnection, originalPrimary);
+    afterSecondariesHaveRestarted(primaryConnection);
 
     // TODO SERVER-109457 Try a scenario where you force election here so that the restarted
     // secondary could become primary while another active secondary has not restarted yet. This